The term of 'quantitative trait nucleotide' (QTN)

The first mention of the term 'QTN' was in the following paper from 1998:

Long A.D., Lyman R.F., Langley C.H. & Mackay T.F. (1998) Two sites in the Delta gene region contribute to naturally occurring variation in bristle number in Drosophila melanogaster. Genetics 149, 999-1017.

The relevant text from the paper (page 1000):

'If a neutral molecular marker (referred to as the marker site) and a site contributing to variation in a quantitative trait (the quantitative trait nucleotide or QTN) are physically close to one another, they are likely to be in linkage disequilibrium.'

Confirmed by Trudy MacKey, in AnGenMap Discussion Mailing List